Laccoptera (Laccopteroidea) nepalensis Boheman, 1855
Cassida quadrimaculata Thunberg, 1789: 86 (LT in Uppsala); Schönherr, 1817: 221; Sekerka, 2008 b: 304 (LT designation), not C. quadrimaculata De Geer, 1775.
Laccoptera quadrimaculata: Boheman, 1856: 156, 1862: 385; Gemminger and Harold, 1876: 3662; Maulik, 1916: 587; 1919: 347 (incl. fig.); Spaeth and Reitter, 1926: 9; Spaeth, 1938 a: 231; Gressitt, 1952: 471, 584 (incl. fig., egg, larva, pupa); Chen and Zia, 1961: 441; Gressitt and Kimoto, 1963: 956 (incl. fig., egg, larva, pupa); Kimoto, 1966: 643; 1991: 26, 1998: 35 (incl. colour photo); Vazirani, 1972: 150; Kimoto and Takizawa, 1973: 180; Takizawa, 1975: 62; 1978: 602; 1980: 22 (larva); 1985 a: 5; 1985 b: 113; 1987: 527; 1988: 10; 1989: 325; Medvedev and Eroshkina, 1982: 107; 1988: 123; Lizhong, 1982: 54; Zaitsev and Medvedev, 1983: 137 (larva); Chen and Zia, 1984: 79; Borowiec, 1985 a: 28; 1990 a: 703; Basu, 1985: 213; Chen et al., 1986: 575, 635 (incl. fig.); Yu, 1986: 681; Basu and Halder, 1987: 238; Verma, 1988: 2; Borowiec and Takizawa, 1991: 638; Zaitsev, 1992: 177 (pupa); Medvedev, 1992: 35, 1997: 265, 2000: 25; Peng and Liu, 1992: 613 (fig.); Mohamedsaid, 1993: 46 (colour photo), 1994: 370, 2000 a: 359; Kimoto and Chu, 1996: 134; Kimoto et al., 1995: 108 (incl. photo); Sprecher-Uebersax, 1997: 161; Sumana Saha and Raychaudhuri, 1997: 83; Medvedev and Sprecher-Uebersax, 1999: 350; Hartmann and Medvedev, 2003: 181.
Laccoptera (Laccoptera) quadrimaculata: Spaeth, 1914 g: 82.
Laccoptera quadrimaculata quadrimaculata: Chen et al., 1986: 576, 635, pl. XIII (incl. colour fig.); Yu, 1986: 681, 2002: 466; Lizhong, 1989: 87.
Laccoptera 4-maculata: Medvedev and Dan, 1982: 96.
Laccoptera Nepalensis Boheman, 1855: 76 (ST! in NRS!, HU!), 1856: 156, 1862: 385; Gemminger and Harold, 1876: 3662; Borowiec, 1996 b: 35, 2001 e: 535 2009 d: 466; Mohamedsaid, 2000 b: 377; Kimoto, 2005: 111 (incl. photo); Sultan et al., 2008: 100 (host plants).
Laccoptera (Laccoptera) nepalensis: Spaeth, 1914 g: 82.
Laccoptera quadrimaculata nepalensis: Zia and Wu, 1981: 521; Yu, 1983: 795, 1986: 681, 1988: 356; Chen et al., 1986: 576, 635.
Laccoptera (Laccopteroidea) nepalensis: Borowiec, 1999 d: 219; Świętojańska, 2001 c: 223 (incl. fig., colour photo); Ghate et al. 2003: 531; Mohamedsaid, 2004: 195; Sekerka, 2004: 159, 2006 c: 99; Borowiec and Sekerka, 2010: 369; Ghosh et al., 2023: 27.
Laccoptera Bohemani Weise, 1910 d: 42 (= Laccoptera chinensis sensu Boheman, 1855; TE in ?); Spaeth, 1913 e: 46, 1914 a: 15.
Laccoptera quadrimaculata ssp. Bohemani: Spaeth, 1914 g: 82.
Laccoptera quadrimaculata ab. bohemani: Chujô, 1934: 153.
Laccoptera quadrimaculata bohemani: Gressitt, 1938 a: 191; 1938 b: 384; 1938 c: 581; 1939: 140.
Laccoptera Thunbergi Spaeth, 1914a: 15 (new name for Cassida quadrimaculata Thunberg, 1789 not De Geer, 1775.
Laccoptera chinensis F.: Boheman, 1855: 71, 1856: 155, 1862: 384; Gemminger and Harold, 1876: 3662 (misidentification).
Laccoptera quatuordecimmaculata Thunberg, 1789 [sic]: V.L. Medvedev, 2002: 36.
Host plant: Convolvulaceae: Calystegia soldanella (Chűjô and Kimoto, 1961); Ipomoea indica (Takizawa, 1975); Ipomoea sp. (Takizawa, 1978 b; Medvedev and Dan, 1982); Ipomoea carnea ssp. fistulosa, I. nil (Sultan et al., 2008); Calystegia sepium, C. silvatica ssp. orientalis, Dinetus racemosus, Ipomoea batatas, Ipomoea purpurea (Yang et al. 2024).
Distr.: quadrimaculata: China; nepalensis: Nepal Indiae orientalis; bohemani: Indien, China. Andaman Is.; Burma; S and E China; India; Indonesia: Java, Sumatra; Japan: Ryukyu Is.; Laos; Malaysia; Nepal; Pakistan; Singapore; Thailand; Vietnam.
figure: habitus in colour (two common forms)
figure: habitus in colour (two rare forms)
figure: adult feeding pattern